https://w3id.org/np/RA84bHmOn6BQnJEYRPAskbEr0S4qdqtwHjEYmt0NMqj9c/Head
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https://w3id.org/np/RA84bHmOn6BQnJEYRPAskbEr0S4qdqtwHjEYmt0NMqj9c/assertion
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
http://purl.org/dc/terms/description
Space: 5883 bird species with a one-to-one BirdLife-BirdTree name match, a tree tip and an AVONET range centroid. Patristic distances were computed on the MCC tree; 1999655 random species pairs were drawn and the Spearman correlation between patristic and great-circle centroid distance was compared with 99 permutations of centroids among species; medians of centroid distance were tabulated for congeneric, confamilial and same-order pairs. Time: PBDB accepted genera with family, order and first-appearance ages; within each order, genus pairs (all pairs, or 500000 random pairs for very large orders) were split into same-family and different-family, and the difference of medians of |dFAD| was compared with 99 permutations of family labels within orders.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
http://www.w3.org/1999/02/22-rdf-syntax-ns#type
https://w3id.org/delphi/ontology/Assessment
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
http://www.w3.org/2000/01/rdf-schema#comment
Measured: 0.026622678561148156 Spearman rho between patristic distance and centroid great-circle distance, bird species pairs (n = 1999655, permutation p = 0.010; permuted rho range -0.0071 to 0.0068). Interpretation: 'Related by descent' is measured for living birds as patristic distance on a dated molecular supertree (Jetz et al. 2012 BirdTree, Hackett backbone MCC tree as distributed with AVONET), a modern reinterpretation of Darwin's classification-based relatedness, and for fossils by shared family versus shared order in PBDB taxonomy, which is closer to Darwin's own usage. 'Nearer in geographic space' is the great-circle distance between AVONET breeding-range centroids. 'Nearer in time' is the absolute difference between first-appearance ages (midpoints of PBDB first-appearance age ranges) of genera. Only two living/fossil systems are tested, not 'all organisms'; birds are chosen because tree, taxonomy and centroids come from a single curated source, and mammals and molluscs because they are the best-sampled terrestrial and marine fossil records. Sampling and taxonomic-practice confounds (geography informing species limits; fossils concentrated in a few basins) are noted as caveats rather than corrected.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
http://www.w3.org/2000/01/rdf-schema#label
Claim test of darwin1859-ch12-18: supports, confidence 0.75
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
http://www.w3.org/2000/01/rdf-schema#seeAlso
https://github.com/fractastical/delphisci/blob/bf2af6ee0a84e23ffa6414ad7a0a50a8c276506e/test-data/claim-tests/darwin-origin/darwin1859-ch12-18/result.json
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
http://www.w3.org/ns/prov#wasAttributedTo
https://w3id.org/delphi/resource/agent/delphisci-claim-tester-claude-fable-5-1
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/admissibleDatasets
Planned datasets: avonet-tobias2022-figshare, pbdb-taxa-first-appearances-mammalia-mollusca
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/assessmentType
claim-test/0.1.0
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/dataKind
biological
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/disconfirmingObservation
The space clause fails if the Spearman correlation is <= 0 or does not exceed the permutation distribution at p <= 0.01, or if the median distances are not strictly ordered by relatedness. The time clause fails if in either group same-family genus pairs are not closer in first-appearance time than different-family same-order pairs, or the difference is not significant at p <= 0.01 against the within-order permutation null. Both failing refutes the claim.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/evidenceAgainst
Family and order assignments in PBDB are of uneven quality, especially for Mollusca; genera with unassigned family or order were dropped.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/evidenceAgainst
Fossil first appearances are truncated by sampling and concentrated in well-sampled basins and intervals; genera in the same family are often described from the same formations, which can make same-family FADs look closer than they are.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/evidenceAgainst
Patristic distance on the BirdTree MCC tree is a model-based divergence estimate, a modern proxy for Darwin's relatedness by descent.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/evidenceAgainst
Pooled permutation nulls for the fossil clause are not centred on zero because orders differ in family structure and age spread; the observed differences are judged against that null, not against zero.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/evidenceAgainst
Range centroids collapse each species to a point; sister species with adjacent but large ranges can appear far apart, which weakens rather than inflates the correlation.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/evidenceAgainst
Space is tested in one class (birds) and time in two fossil groups at genus level; the claim is stated for all organisms, living and fossil.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/evidenceAgainst
Species limits in birds have historically been drawn partly on geography (allopatric forms), which can tighten the congeneric-pair pattern by construction.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/evidenceAgainst
The overall Spearman correlation between patristic and geographic distance is small (about 0.03) although far outside the permutation null; the relationship is concentrated at shallow phylogenetic depth (congeneric and confamilial pairs) and saturates at the order level, where median centroid distance is no smaller within than between orders.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/expectedObservation
Space (all extant birds with a range centroid in AVONET and a tip in the BirdTree Hackett MCC tree supplied with AVONET): (S1) the Spearman correlation between patristic distance and great-circle centroid distance across a fixed random sample of 2,000,000 species pairs is positive, and exceeds all of 99 permutations in which centroids are shuffled among species (p <= 0.01); (S2) median centroid distance is strictly ordered congeneric pairs < confamilial non-congeneric pairs < same-order different-family pairs. Time (Paleobiology Database genera of Mammalia and of Mollusca with first-appearance ages and family/order assignments): (T1) in each of the two groups, the median absolute difference in first-appearance midpoint age between pairs of genera in the same family is smaller than between pairs of genera in different families of the same order, and this difference exceeds all of 99 permutations of family labels within orders (p <= 0.01). The space clause passes if S1 and S2 hold; the time clause passes if T1 holds in both groups. Verdict 'supports' if both clauses pass, 'mixed' if one passes, 'refutes' if neither.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/failCriterion
The space clause fails if the Spearman correlation is <= 0 or does not exceed the permutation distribution at p <= 0.01, or if the median distances are not strictly ordered by relatedness. The time clause fails if in either group same-family genus pairs are not closer in first-appearance time than different-family same-order pairs, or the difference is not significant at p <= 0.01 against the within-order permutation null. Both failing refutes the claim.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/hasConfidence
0.75
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/hasEvidence
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-1
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/hasEvidence
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-2
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/hasVerdict
supports
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/hypothesis
If closer relatives are generally nearer in space and time, then (space, living forms) phylogenetic distance between bird species should correlate positively with the geographic distance between their range centroids, with congeneric pairs closer than confamilial pairs and those closer than pairs from different families of the same order; and (time, fossil forms) genera in the same family should have first appearances closer in geological time than genera in different families of the same order.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/inferenceMethod
Space: 5883 bird species with a one-to-one BirdLife-BirdTree name match, a tree tip and an AVONET range centroid. Patristic distances were computed on the MCC tree; 1999655 random species pairs were drawn and the Spearman correlation between patristic and great-circle centroid distance was compared with 99 permutations of centroids among species; medians of centroid distance were tabulated for congeneric, confamilial and same-order pairs. Time: PBDB accepted genera with family, order and first-appearance ages; within each order, genus pairs (all pairs, or 500000 random pairs for very large orders) were split into same-family and different-family, and the difference of medians of |dFAD| was compared with 99 permutations of family labels within orders.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/passCriterion
Space (all extant birds with a range centroid in AVONET and a tip in the BirdTree Hackett MCC tree supplied with AVONET): (S1) the Spearman correlation between patristic distance and great-circle centroid distance across a fixed random sample of 2,000,000 species pairs is positive, and exceeds all of 99 permutations in which centroids are shuffled among species (p <= 0.01); (S2) median centroid distance is strictly ordered congeneric pairs < confamilial non-congeneric pairs < same-order different-family pairs. Time (Paleobiology Database genera of Mammalia and of Mollusca with first-appearance ages and family/order assignments): (T1) in each of the two groups, the median absolute difference in first-appearance midpoint age between pairs of genera in the same family is smaller than between pairs of genera in different families of the same order, and this difference exceeds all of 99 permutations of family labels within orders (p <= 0.01). The space clause passes if S1 and S2 hold; the time clause passes if T1 holds in both groups. Verdict 'supports' if both clauses pass, 'mixed' if one passes, 'refutes' if neither.
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/ontology/preregistered
false
https://w3id.org/delphi/resource/claim/darwin1859-ch12-18
https://w3id.org/delphi/ontology/hasAssessment
https://w3id.org/delphi/resource/assessment/darwin1859-ch12-18--claim-test
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-1
http://purl.org/dc/terms/description
phylogenetic distance, range centroids and taxonomy of extant birds
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-1
http://purl.org/dc/terms/license
CC BY 4.0
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-1
http://www.w3.org/1999/02/22-rdf-syntax-ns#type
https://w3id.org/delphi/ontology/EvidenceItem
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-1
http://www.w3.org/2000/01/rdf-schema#label
Dataset: AVONET ELEData.zip: TraitData/AVONET1_BirdLife.csv (range centroids, taxonomy), PhylogeneticData/HackettStage1_0001_1000_MCCTreeTargetHeights.nex (BirdTree Hackett MCC tree, 6670 species) and BirdLife-BirdTree crosswalk.csv
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-1
http://www.w3.org/ns/prov#atLocation
https://ndownloader.figshare.com/files/38429873
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-1
http://www.w3.org/ns/prov#wasDerivedFrom
https://figshare.com/articles/dataset/AVONET_morphological_ecological_and_geographical_data_for_all_birds_Tobias_et_al_2021_Ecology_Letters_/16586228
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-1
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https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-2
http://purl.org/dc/terms/description
first-appearance ages and family/order of fossil genera
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-2
http://purl.org/dc/terms/license
CC BY 4.0
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-2
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https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-2
http://www.w3.org/2000/01/rdf-schema#label
Dataset: Paleobiology Database taxa lists (genus rank) with first/last appearance ages and classification for Mammalia and Mollusca
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-2
http://www.w3.org/ns/prov#atLocation
https://paleobiodb.org/data1.2/taxa/list.json?base_name=Mammalia&rank=genus&show=app%2Cclass&vocab=pbdb
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-2
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https://paleobiodb.org/data1.2/taxa/list.json?base_name=Mollusca&rank=genus&show=app%2Cclass&vocab=pbdb
https://w3id.org/delphi/resource/evidence/darwin1859-ch12-18--claim-test--dataset-2
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delphisci claim tester (claude-fable-5.1; cursor/claim-tests-darwin-batch-0e75)
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2026-11-08T16:24:31.858Z
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Claim test of darwin1859-ch12-18: supports, confidence 0.75
https://w3id.org/np/RA84bHmOn6BQnJEYRPAskbEr0S4qdqtwHjEYmt0NMqj9c
https://w3id.org/delphi/ontology/domain
https://w3id.org/delphi/resource/domain/darwin-origin-1859
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RSA
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